A greater range of shade-tolerance niches in nutrient-rich forests: an explanation for positive richness–productivity relationships?

Investigating if a wider range of growth rates and shade tolerances are found on nutrient-rich soils, because such soils not only support fast-growing species with high metabolic rates, but also species capable of tolerating the very deep shade cast by forest canopies growing where nutrients are plentiful.

Coomes, D.A.; Kunstler, G.; Canham, C.D.; Wright, E.